Costa, W. J. E. M. 1998. Phylogeny and classification of Rivulidae revisited: Origin and evolution of annualism and miniaturization in rivulid fishes (Cyprinodontiformes: Aplocheiloidei). Journal of Comparative Biology 3 (1): 33-92, 37 figs. [description: p. 77]

Type species: Pterolebias zonatus Myers, 1935, by original designation.

Gnatholebias Costa, 1998.